• Home
  • Latest
  • Fortune 500
  • Finance
  • Tech
  • Leadership
  • Lifestyle
  • Rankings
  • Multimedia
EnvironmentAnimals

Gramma, Galapagos tortoise and oldest resident of San Diego Zoo, dies at (probably) 141 years old

By
Jaimie Ding
Jaimie Ding
and
The Associated Press
The Associated Press
Down Arrow Button Icon
By
Jaimie Ding
Jaimie Ding
and
The Associated Press
The Associated Press
Down Arrow Button Icon
November 25, 2025, 11:06 AM ET
Gramma
This photo provided by the San Diego Zoo Wildlife Alliance shows Gramma, a Galapagos tortoise and the oldest animal at the San Diego Zoo, eating a banana stalk at the San Diego Zoo in San Diego, May 17, 2023. San Diego Zoo Wildlife Alliance via AP

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Recommended Video

Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died Nov. 20.

It’s not clear exactly when the tortoise arrived at the San Diego Zoo, but zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galapagos tortoises.

As the world changed around her, she delighted visitors with her sweet, shy personality. She lived through two World Wars and 20 U.S. presidents.

Her care specialists affectionately called her “the Queen of the Zoo.” She was suffering from bone conditions related to her old age that progressed recently before she was euthanized, the zoo said.

Many visitors commented on social media about getting to first visit Gramma when they were young, and being able to come back years later with their kids.

Cristina Park, 69, said one of her earliest memories from her childhood was going to the San Diego Zoo when she was 3 or 4 years old and riding on the back of a tortoise. That’s no longer allowed, but the experience inspired her to keep a small desert tortoise as a pet and learn more about tortoise conservation.

“Just how amazing it is that they managed to live through so much,” Park said. “And yet they’re still there.”

Galapagos tortoises can live for over 100 years in the wild, and close to double that in captivity.

The oldest known Galapagos tortoise was named Harriet, who lived at the Australia Zoo until the age of 175. She was collected from the Galapagos Islands in 1835, when she was just the size of a dinner plate, according to the zoo. This means that she hatched somewhere around 1830, and she died in 2006.

Galapagos tortoises include 15 subspecies of tortoises from the islands, three of which were deemed extinct. The rest are all vulnerable or critically endangered, according to the International Union for Conservation of Nature.

Concerted efforts have been made to breed these tortoises in captivity over the past several decades, with more than 10,000 juveniles released to the wild since 1965, according to the Galapagos Conservancy. Some subspecies have been brought back from the brink of extinction.

In April, four baby Galapagos tortoises were born at the Philadelphia Zoo to first-time parents that were roughly 100 years old, a first in the zoo’s history. In June, Zoo Miami resident and Galapagos tortoise Goliath became a first-time father at the age of 135.

The Fortune 500 Innovation Forum will convene Fortune 500 executives, U.S. policy officials, top founders, and thought leaders to help define what’s next for the American economy, Nov. 16-17 in Detroit. Apply here.
About the Authors
By Jaimie Ding
See full bioRight Arrow Button Icon
By The Associated Press
See full bioRight Arrow Button Icon

Latest in Environment

Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025

Most Popular

Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Finance
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut labore et dolore magna aliqua. Ut enim ad minim veniam
By Fortune Editors
October 20, 2025
Fortune Secondary Logo
Rankings
  • 100 Best Companies
  • Fortune 500
  • Global 500
  • Fortune 500 Europe
  • Most Powerful Women
  • Future 50
  • World’s Most Admired Companies
  • See All Rankings
Sections
  • Finance
  • Fortune Crypto
  • Features
  • Leadership
  • Health
  • Commentary
  • Success
  • Retail
  • Mpw
  • Tech
  • Lifestyle
  • CEO Initiative
  • Asia
  • Politics
  • Conferences
  • Europe
  • Newsletters
  • Personal Finance
  • Environment
  • Magazine
  • Education
Customer Support
  • Frequently Asked Questions
  • Customer Service Portal
  • Privacy Policy
  • Terms Of Use
  • Single Issues For Purchase
  • International Print
Commercial Services
  • Advertising
  • Fortune Brand Studio
  • Fortune Analytics
  • Fortune Conferences
  • Business Development
  • Group Subscriptions
About Us
  • About Us
  • Editorial Calendar
  • Press Center
  • Work At Fortune
  • Diversity And Inclusion
  • Terms And Conditions
  • Site Map
  • About Us
  • Editorial Calendar
  • Press Center
  • Work At Fortune
  • Diversity And Inclusion
  • Terms And Conditions
  • Site Map
  • Facebook icon
  • Twitter icon
  • LinkedIn icon
  • Instagram icon
  • Pinterest icon

Latest in Environment

david-f
CommentaryVenture Capital
Europe has survived 3 energy shocks in 4 years. The only way out is to stop buying power from its enemies
By David FrykmanMarch 25, 2026
4 hours ago
Farmer standing in front a soybean farm
Economyfertilizer
Soaring fertilizer prices could pressure a U.S. agricultural industry that supports 50 million jobs and over $10 trillion in output
By Tristan BoveMarch 24, 2026
17 hours ago
fauci
CommentaryCOVID-19 vaccines
How COVID turned America against science — and what it will take to win it back
By David Blumenthal and James A. MoroneMarch 24, 2026
1 day ago
EnergyEnergy Industry
Trump despises wind farms so much he’s paying a French energy giant $1 billion to stop building them and invest in natural gas instead
By Jordan BlumMarch 23, 2026
2 days ago
HealthHealth
Forest ‘bathing’ can reduce stress, improve mood, lower blood pressure and boost the immune system. Here’s how it’s done
By Allen Breed and The Associated PressMarch 22, 2026
3 days ago
Stephan Winkelmann looks forward. He is wearing a suit.
EnergyLuxury
Lamborghini CEO says ‘disappointing’ EV charging infrastructure contributed to no demand for the luxury automaker’s all-electric line
By Sasha RogelbergMarch 20, 2026
5 days ago

Most Popular

Magazine
The youngest-ever female CEO of a Fortune 500 company is fighting Trump's cuts to keep Medicaid strong
By Fortune EditorsMarch 24, 2026
1 day ago
Commentary
The Treasury just declared the U.S. insolvent. The media missed it
By Fortune EditorsMarch 23, 2026
2 days ago
Economy
It took 200 years for national debt to hit $1 trillion. Annual interest alone now exceeds that—a 'crushing legacy we must reverse,' says budget chair
By Fortune EditorsMarch 23, 2026
2 days ago
Energy
Nobel laureate Paul Krugman calls it 'treason': $580 million in suspicious oil futures traded minutes before Trump's Iran reversal
By Fortune EditorsMarch 24, 2026
17 hours ago
Success
Palantir’s billionaire CEO says only two kinds of people will succeed in the AI era: trade workers — ‘or you’re neurodivergent’
By Fortune EditorsMarch 24, 2026
21 hours ago
Personal Finance
Current price of oil as of March 24, 2026
By Fortune EditorsMarch 24, 2026
24 hours ago

© 2026 Fortune Media IP Limited. All Rights Reserved. Use of this site constitutes acceptance of our Terms of Use and Privacy Policy | CA Notice at Collection and Privacy Notice | Do Not Sell/Share My Personal Information
FORTUNE is a trademark of Fortune Media IP Limited, registered in the U.S. and other countries. FORTUNE may receive compensation for some links to products and services on this website. Offers may be subject to change without notice.